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2023.  Ghose*, S. L., T. A. Yap, A. Q. Byrne, H. Sulaeman, E. B. Rosenblum, W. Bauer, A. Chan-Alvarado, S. Chaukulkar, E. Greenbaum, K. Lutz, D. McAloose, A. Moyer, E. Parra, D. M. Portik, H. Rockney, A. G. Zink, D. C. Blackburn, and V. T. Vredenburg. Continent-wide recent emergence of a global pathogen in African amphibians. Frontiers in Conservation Science 4:1069490.

2023.  Das, S., E. Greenbaum, S. Meiri, A. M. Bauer, F. T. Burbrink, C. J. Raxworthy, J. L. Weinell, R. M. Brown, J. Brecko, O. S. G. Pauwels, N. Rabibisoa, A. P. Raselimanana, and J. Merilä. Ultraconserved elements-based phylogenomic systematics of the snake superfamily Elapoidea, with the description of a new Afro-Asian family. Molecular Phylogenetics and Evolution 180:107700.

2022. Greenbaum, E., D. M. Portik, K. E. Allen*, E. R. Vaughan, G. Badjedjea, M. F. Barej, M. Behangana, N. Conkey**, B. Dumbo, L. N. Gonwouo, M. Hirschfeld, D. F. Hughes, F. Igunzi, C. Kusamba, W. Lukwago, F. M. Masudi, J. Penner, J. Reyes**, M. -O. Rödel, C. E. Roelke, S. Romero**, and J. M. Dehling. Systematics of the Central African spiny reed frog Afrixalus laevis (Anura: Hyperoliidae), with the description of two new species from the Albertine Rift. Zootaxa 5174:201–232.

2022.  Keating, S. E.*, E. Greenbaum, J. D. Johnson, and T. Gamble. Identification of a cis-sex chromosome transition in banded geckos (Coleonyx, Eublepharidae, Gekkota). Journal of Evolutionary Biology 35:1675–1682. 

2022. Jaynes*, K. E., E. A. Myers, V. Gvoždík, D. C. Blackburn, D. M. Portik, E. Greenbaum, G. F. M. Jongsma, M. -O. Rödel, G. Badjedjea, A. Bamba-Kaya, N. L. Baptista, J. B. Akuboy, R. Ernst, M. T. Kouete, C. Kusamba, F. M. Masudi, P. J. McLaughlin, L. M. Nneji, A. B. Onadeko, J. Penner, P. Vaz Pinto, B. L. Stuart, E. Tobi, A. -G. Zassi-Boulou, A. D. Leaché, M. K. Fujita, and R. C. Bell. Giant tree frog diversification in West and Central Africa: Isolation by physical barriers, climate, and reproductive traits. Molecular Ecology 31: 3979–3998.

2021. Teshera*, M. S., R. W. Clark, A. E. Wagler, and E. Greenbaum. Foraging and scavenging behaviour of the pairie rattlesnake (Crotalus viridis): no evidence that envenomation cues facilitate kleptoparasitism of struck prey. Amphibia-Reptilia 42:491–501.

2021. Goldberg, S. R., C. R. Bursey, F. M. Mali, C. Kusamba, A. J. Robbins*, and E. Greenbaum. Gastrointestinal helminths in Amietia sp. (Anura: Pyxicephalidae) from the Albertine Rift of Central Africa. Helminthologia 58:328–332.

2021. Engel, M. S., L. M. P. Ceríaco, G. M. Daniel, P. M. Dellapé, I. Löbl, M. Marinov, R. Reis, M. T. Young, A. Dubois, I. Agarwal, P. Lehmann Albornoz, M. Alvarado, N. Alvarez, F. Andreone, K. Araujo-Vieira, J. S. Ascher, D. Baêta, D. Baldo, S. A. Bandeira, P. Barden, D. A. Barrasso, L. Bendifallah, F. A. Bockmann, W. Böhme, A. Borkent, C. R. F. Brandão, S. D. Busack, S. M. Bybee, A. Channing, S. Chatzimanolis, M. J. M. Christenhusz, J. V. Crisci, G. d’Elía, L. M. Da Costa, S. R. Davis, C. Alberto S. de Lucena, T. Deuve, S. Fernandes Elizalde, J. Faivovich, H. Farooq, A. W. Ferguson, S. Gippoliti, F. M. P. Gonçalves, V. H. Gonzalez, E. Greenbaum, I. A. Hinojosa-Díaz, I. Ineich, J. Jiang, S. Kahono, A. B. Kury, V. Lohrmann, P. H. F. Lucinda, J. D. Lynch, V. Malécot, M. P. Marques, J. W. M. Marris, R. C. McKellar, L. F. Mendes, S. S. Nihei, K. Nishikawa, A. Ohler, V. G. D. Orrico, H. Ota, J. Paiva, D. Parrinha, O. S. G. Pauwels, M. O. Pereyra, L. B. Pestana, P. D. P. Pinheiro, L. Prendini, J. Prokop, C. Rasmussen, M. -O. Rödel, M. Trefaut Rodrigues, S. M. Rodríguez, H. Salatnaya, í. Sampaio, A. Sánchez-García, M. A. Shebl, B. S. Santos, M. M. Solórzano-Kraemer, A. C. A. Sousa, P. Stoev, P. Teta, J. -F. Trape, C. Van-Dúnem Dos Santos, K. Vasudevan, C. J. Vink, G. Vogel, P. Wagner, T. Wappler, J. L. Ware, S. Wedmann, C. Kusamba Zacharie, and T. Ziegler. The taxonomic impediment: A shortage of taxonomists, not the lack of technical approaches. Zoological Journal of the Linnean Society 193:381–387.

2021. Blackburn, D. C., S. V. Nielsen, S. L. Ghose, M. Burger, L. N. Gonwouo, E. Greenbaum, V. Gvoždík, M. Hirschfeld, M. T. Kouete, C. Kusamba, D. Lawson, P. J. McLaughlin, A. -G. Zassi-Boulou, and M. -O. Rödel. Phylogeny of African long-fingered frogs (Arthroleptidae: Cardioglossa) reveals recent allopatric divergences in coloration. Ichthyology & Herpetology 109:728–742.

2021. Goldberg, S. R., C. R. Bursey, C. Kusamba, F. M. Mali, and E. Greenbaum. Helminths from three common species of African toads (Anura: Bufonidae: Sclerophrys). Alytes 38:49–57.

2021. Ceríaco, L. M. P., A. M. Bauer, C. Kusamba, I. Agarwal, and E. Greenbaum. A new species of ground dwelling Hemidactylus (Squamata: Gekkonidae) from southwestern Democratic Republic of the Congo. Journal of Herpetology 55:105–111.

2021. Allen*, K. E., E. Greenbaum, P. M. Hime, W. P. Tapondjou N*, V. T. Sterkhova**, C. Kusamba, M. -O. Rödel, J. Penner, A. Towsend Peterson, and R. M. Brown. Rivers, not refugia, drove diversification in arboreal, sub-Saharan African snakes. Ecology and Evolution 11:6133–6152.

2021. Greenbaum, E., K. E. Allen*, E. R. Vaughan*, O. S. G. Pauwels, V. Wallach, C. Kusamba, W. M. Muninga, M. M. Aristote, F. M. M. Mali*, G. Badjedjea, J. Penner, M. -O. Rödel, J. Rivera**, V. Sterkhova**, G. Johnson**, W. P. Tapondjou N*, and R. M. Brown. Night stalkers from above: A monograph of Toxicodryas tree snakes (Squamata: Colubridae) with descriptions of two new cryptic species from Central Africa. Zootaxa 4965:001–044.

2020. Zimkus, B. M., V. Baláž, A. M. Belasen, R. C. Bell, A. Channing, J. Doumbia, E. B. Fokam, L. N. Gonwouo, E. Greenbaum, V. Gvoždík, M. Hirschfeld, K. Jackson, T. Y. James, C. Kusamba, J. G. Larson, L. -B. Mavoungou, M. -O. Rödel, A. -G. Zassi-Boulou, and J. Penner. Chytrid fungus (Batrachochytrium dendrobatidis) in African amphibians: A continental analysis of occurrences and modeling of its potential distribution. Herpetologica 76:201–215.

2020. Greenwood, L., S. P. Loader, L. Lawson, E. Greenbaum, and B. M. Zimkus. A new species of Phrynobatrachus(Amphibia: Anura: Phrynobatrachidae) from the northern mountains of Tanzania. Journal of Natural History 54:63–85.

2020. Engelbrecht, H., W. R. Branch, E. Greenbaum, M. Burger, W. Conradie, and K. A. Tolley. African herald snakes, Crotaphopeltis, show population structure for a widespread generalist but deep genetic divergence for forest specialists. Journal of Zoological Systematics and Evolutionary Research 58:1220–1233.

2020. Cauret*, C. M. S., M. -T. T. Gansauge, A. S. Tupper*, B. L. S. Furman, M. Knytl, X. Song*, E. Greenbaum, M. Meyer, and B. J. Evans. Developmental systems drift and the drivers of sex chromosome evolution. Molecular Biology and Evolution 37:799–810.

2019. Keates*, C., W. Conradie, E. Greenbaum, and S. Edwards. Genetic structuring of the widespread genus Psammophylax (Fitzinger 1843), with the description of a new genus and a new species. Journal of Zoological Systematics and Evolutionary Research 57:1039–1066.

2019. Leaché, A. D., D. M. Portik, D. Rivera, M.-O. Rödel, J. Penner, V. Gvoždík, E. Greenbaum, C. Ofori-Boateng, M. Burger, E. A. Eniang, R. C. Bell, and M. K. Fujita. Exploring rainforest diversification using demographic model testing in the African foam-nest treefrog (Chiromantis rufescens). Journal of Biogeography 46:2706–2721.

2019. Vaughan*, E. R., M. S. Teshera*, C. Kusamba, T. R. Edmonston**, and E. Greenbaum. A remarkable example of Batesian mimicry of Gaboon Vipers (Reptilia: Viperidae: Bitis gabonica) by Congolese Giant Toads (Amphibia: Bufonidae: Sclerophrys channingi). Journal of Natural History 53:1853–1871.

2019. Allen, K. E.*, W. P. Tapondjou N, E. Greenbaum, L. J. Welton, and A. M. Bauer. High levels of hidden phylogenetic structure and diversity within Central and West African Trachylepis skinks. Salamandra 55:231–241.

2019. Greenbaum, E., and M. V. Zhuang. Robert Gravem Webb (1927–2018): Specialist on trionychid turtles and Mexican herpetology. Herpetological Review 50:656–661.

2019. Evans, B. J., M. -T. T. Gansauge, E. L. Stanley, B. L. S. Furman, C. M. Cauret*, C. Ofori-Boateng, V. Gvoždík, J. W. Streicher, E. Greenbaum, R. C. Tinsley, M. Meyer, and D. C. Blackburn. Xenopus fraseri: Mr. Fraser, where did your frog come from? PLoS ONE 14:e0220892

2019. Portik, D. M., R. C. Bell, D. C. Blackburn, A. M. Bauer, C. D. Barratt, W. R. Branch, M. Burger, A. Channing, T. J. Colston, W. Conradie, J. Maximillian Dehling, R. C. Drewes, R. Ernst, E. Greenbaum, V. Gvoždík, J. Harvey, A. Hillers, M. Hirschfeld, G. F. M. Jongsma, J. Kielgast, M. T. Kouete, L. Lawson, A. D. Leaché, S. P. Loader, S. Lötters, A. van der Meijden, M. Menegon, S. Müller, Z. T. Nagy, C. Ofori-Boateng, A. Ohler, T. J. Papenfuss, D. Rössler, U. Sinsch, M. -O. Rödel, M. Veith, J. Vindum, A. -G. Zassi-Boulou, and J. A. McGuire. Sexual dichromatism drives diversification within a major radiation of African amphibians. Systematic Biology 68:859–875.

2019. Portillo*, F., E. L. Stanley, W. R. Branch, W. Conradie, M. -O. Rödel, J. Penner, M. F. Barej, C. Kusamba, W. M. Muninga, M. M. Aristote, A. M. Bauer, J. -F. Trape, Z. T. Nagy, P. Carlino, O. S. G. Pauwels, M. Menegon, I. Ineich, M. Burger, A.-G. Zassi-Boulou, T. Mazuch, K. Jackson, D. F. Hughes*, M. Behangana, and E. Greenbaum. Evolutionary history of burrowing asps (Squamata: Lamprophiidae: Atractaspidinae) with emphasis on fang evolution and prey selection. PLoS ONE 14:e0214889.

2019. Portillo*, F., W. R. Branch, C. R. Tilbury, Z. T. Nagy, D. F. Hughes*, C. Kusamba, W. M. Muninga, M. M. Aristote, M. Behangana, and E. Greenbaum. A cryptic new species of Polemon (Squamata: Lamprophiidae, Aparallactinae) from the miombo woodlands of Central and East Africa. Copeia 107:22–35.

2019. Engelbrecht*, H. M., W. R. Branch, E. Greenbaum, G. J. Alexander, K. Jackson, M. Burger, W. Conradie, C. Kusamba, A. -G. Zassi-Boulou, and K. A. Tolley. Diversifying into the branches: Species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution 130:357–365.

2018. Bursey, C. R., S. R. Goldberg, C. Kusamba, and E. Greenbaum. New species of Falcaustra (Nematoda: Kathlaniidae) and other helminths in Amietia (Anura: Pyxicephalidae) from the Albertine Rift of Central Africa. Comparative Parasitology 85:177–181.

2018 Greenbaum, E., S. A. Dowell Beer, P. Wagner, C. G. Anderson*, C. O. Villanueva**, P. Malonza, C. Kusamba, W. M. Muninga, M. M. Aristote, and W. R. Branch. Phylogeography of Jackson’s Forest Lizard Adolfus jacksoni (Sauria: Lacertidae) reveals cryptic diversity in the highlands of East Africa. Herpetological Monographs 32:51–68.

2018. Wüster, W., L. Chirio, J. -F. Trape, I. Ineich, K. Jackson, E. Greenbaum, C. Kusamba, C. Barron**, Z. Nagy, R. Storey, C. Hall, C. Wüster, A. Barlow, and D. G. Broadley. Integration of nuclear and mitochondrial gene sequences and morphology reveal unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae). Zootaxa 4455:68–98.

2018. Mayfield, T. J., J. T. Olimpo, K. W. Floyd, and E. Greenbaum. Collaborative position posters develop students’ ability to communicate science to the public. Journal of Microbiology and Biology Education 19:doi:10.1128/jmbe.v19i1.1442 (3 pp).

2018. Wagner, P., E. Greenbaum, A. M. Bauer, C. Kusamba, and A. Leachè. Lifting the blue-headed veil—Integrative taxonomy of the cryptic Acanthocercus atricollis complex (Squamata: Agamidae). Journal of Natural History 52:771–817.

2018. Portillo*, F., W. R. Branch, W. Conradie, M.-O. Rödel, J. Penner, M. F. Barej, C. Kusamba, W. M. Muninga, M. M. Aristote, A. M. Bauer, J.-F. Trape, Z. T. Nagy, P. Carlino, O. S. G. Pauwels, M. Menegon, M. Burger, T. Mazuch, K. Jackson, D. F. Hughes*, M. Behangana, A.-G. Zassi-Boulou, and E. Greenbaum. Phylogeny and biogeography of the African burrowing snake subfamily Aparallactinae (Squamata: Lamprophiidae). Molecular Phylogenetics and Evolution 127:288–303.

2018. Broadley, D. G., K. A. Tolley, W. Conradie, S. Casola, J. -F. Trape, M. Burger, C. Kusamba, and E. Greenbaum. A phylogeny and genus-level revision of the African File Snakes Gonionotophis Boulenger (Squamata: Lamprophiidae). African Journal of Herpetology 67:43–60.

2018. Hughes, D. F.*, K. A. Tolley, M. Behangana, W. Lukwago, M. Menegon, J. M. Dehling, J. Stipala, C. R. Tilbury, A. M. Khan, C. Kusamba, and E. Greenbaum. Cryptic diversity in Rhampholeon boulengeri (Sauria: Chamaeleonidae), a pygmy chameleon from the Albertine Rift biodiversity hotspot. Molecular Phylogenetics and Evolution 122:125–141.

2018. Jongsma*, G. F. M., M. F. Barej*, C. D. Barratt, M. Burger, W. Conradie, R. Ernst, E. Greenbaum, M. Hirschfeld, A. D. Leaché, J. Penner, D. M. Portik, A. -G. Zassi-Boulou, M. -O. Rödel, and D. C. Blackburn. Diversity and biogeography of frogs in the genus Amnirana (Family Ranidae) across sub-Saharan Africa. Molecular Phylogenetics and Evolution 120:274–285.

2017. Bell, R. C., J. L. Parra, G. Badjedjea*, M. F. Barej, D. C. Blackburn, M. Burger, A. Channing, J. M. Dehling, E. Greenbaum, V. Gvoždík, J. Kielgast, C. Kusamba, S. Lötters, P. J. McLaughlin, Z. T. Nagy, M. -O. Rödel, D. M. Portik, B. L. Stuart, J. VanDerWal, and K. R. Zamudio. Idiosyncratic responses to climate-driven forest fragmentation and marine incursions in reed frogs from Central Africa and the Gulf of Guinea Islands. Molecular Ecology 26:5223–5244.

2017. Hughes*, D. F., C. Kusamba, M. Behangana, and E. Greenbaum. Integrative taxonomy of the Central African forest chameleon, Kinyongia adolfifriderici (Sauria: Chamaeleonidae), reveals underestimated species diversity in the Albertine Rift. Zoological Journal of the Linnean Society 181:400–438.

2017. Gamble, T., E. Greenbaum, T. R. Jackman, A. P. Russell, and A. M. Bauer. Repeated evolution of digital adhesion in geckos, a reply to Harrington and Reeder. Journal of Evolutionary Biology 30:1429–1436.

2017. Agarwal, I., S. Biswas, A. M. Bauer, E. Greenbaum, T. Jackman, A. de Silva, and S. Batuwita. Cryptic species, taxonomic inflation, or a bit of both? New species phenomenon in Sri Lanka as suggested by a phylogeny of dwarf geckos (Reptilia, Squamata, Gekkonidae, Cnemaspis). Systematics and Biodiversity 15:427–439.

2017. Leaché, A. D., J. A. Grummer, M. Miller, S. Krishnan, M. K. Fujita, W. Böhme, A. Schmitz, M. LeBreton, I. Ineich, L. Chirio, C. Ofori-Boateng, E. A. Eniang, E. Greenbaum, M. -O. Rödel, and P. Wagner. Bayesian inference of species diffusion in the West African Agama agama species group (Reptilia, Agamidae). Systematics and Biodiversity 15:192–203.

2017. Zimkus, B. M., L. P. Lawson, M. F. Barej, C. D. Barratt, A. Channing, K. M. Dash*, J. Maximilian Dehling, L. Du Preez, P.-S. Gehring, E. Greenbaum, V. Gvoždík, J. Harvey, J. Kielgast, C. Kusamba, Z. T. Nagy, M. Pabijan, J. Penner, M.-O. Rödel, M. Vences, and S. Lötters. Leapfrogging into new territory: How Mascarene ridged frogs diversified across Africa and Madagascar to maintain their ecological niche. Molecular Phylogenetics and Evolution 106:254–269.

2016. Goldberg, S. R., C. R. Bursey, and E. Greenbaum. Nematodes of Leptopelis christyi and Leptopelis karissimbensis (Anura: Arthroleptidae), from the Democratic Republic of the Congo. Alytes 33:21–24.

2016. Blackburn, D. C., E. Greenbaum, M. Fabrezi, D. Meirte, A. J. Plumptre, and E. L. Stanley. The distribution of the Bururi Long-Fingered Frog (Cardioglossa cyaneospila, Family Arthroleptidae), a poorly known Albertine Rift endemic. Zootaxa 4170:355–364.

2016. Hughes*, D. F., E. M. Walker*, P. M. Gignac, A. Martinez*, K. Negishi*, C. L. Lieb, E. Greenbaum, and A. M. Khan. Rescuing perishable neuroanatomical information from a threatened biodiversity hotspot: Remote field methods for brain tissue preservation validated by cytoarchitectonic analysis, immunohistochemistry, and x-ray microcomputed tomography. PLoS ONE 11:e0155824.

2016. Medina*, M. F., A. M. Bauer, W. R. Branch, A. Schmitz, W. Conradie, Z. T. Nagy, T. J. Hibbitts, R. Ernst, D. M. Portik*, S. V. Nielsen*, C. Kusamba, T. J. Colston*, M. Behangana, M. -O. Rödel, and E. Greenbaum. Molecular phylogeny of Panaspis and Afroablepharus skinks (Squamata: Scincidae) in the savannas of sub-Saharan Africa. Molecular Phylogenetics and Evolution 100:409–423.

2016. Larson*, T., D. Castro**, M. Behangana, and E. Greenbaum. Evolutionary history of the river frog genus Amietia (Anura: Pyxicephalidae) reveals extensive diversification in Central African highlands. Molecular Phylogenetics and Evolution 99:168–181.

2016. Dowell*, S. A., D. M. Portik*, V. de Buffrénil, I. Ineich, E. Greenbaum, S. -O. Kolokotronis, and E. R. Hekkala. Molecular data from contemporary and historical collections reveal a complex story of cryptic diversification in the Varanus (Polydaedalus) niloticus Species Group. Molecular Phylogenetics and Evolution 94:591–604.

2015. Seimon, T., S. Ayebare*, R. Sekisambu, E. Muhindo, G. Mitamba, E. Greenbaum, M. Menegon, F. Pupin*, D. McAloose, A. Ammazzalorso, D. Meirte, W. Lukwago*, M. Behangana, D. Kujirakwinja, A. Seimon, and A. J. Plumptre. Assessing the threat of amphibian chytrid fungus in the Albertine Rift: Past, present and future. PLoS ONE 10:e0145841.

2015. Evans, B. J., T. F. Carter, E. Greenbaum, V. Gvoždík, D. B. Kelley, P. J. McLaughlin, O. S. G. Pauwels, D. M. Portik*, E. L. Stanley, R. C. Tinsley, M. L. Tobias, and D. C. Blackburn. Genetics, morphology, acoustical data, and historical records distinguish six new polyploid species of African clawed frog (Xenopus, Pipidae) from West and Central Africa. PLoS ONE 10:e0142823.

2015. Lehr, E., J. Maximilian Dehling, E. Greenbaum, and U. Sinsch. Embryogenesis and tadpole description of Hyperolius castaneus Ahl, 1931 and H. jackie Dehling, 2012 (Anura: Hyperoliidae) in montane bog pools. Zookeys 546:125–152.

2015. Hirschfeld, M*, D. C. Blackburn, M. Burger, E. Greenbaum, A. -G. Zassi-Boulou, and M. -O. Rödel. Two new species of Cardioglossa (Anura: Arthroleptidae) from the Central African rainforests. African Journal of Herpetology. 64:81–102.

2015. Gamble, T., E. Greenbaum, T. R. Jackman, and A. M. Bauer. Into the light: Diurnality has evolved multiple times in geckos. Biological Journal of the Linnean Society 115:896–910.

2015. Greenbaum, E., J. Meece, K. Reed, and C. Kusamba. Extensive occurrence of the amphibian chytrid fungus in the Albertine Rift, a Central African amphibian hotspot. The Herpetological Journal 25:91–100.

2015. Greenbaum, E., F. Portillo*, K. Jackson, and C. Kusamba. A phylogeny of Central African Boaedon (Serpentes: Lamprophiidae), with the description of a new species from the Albertine Rift. African Journal of Herpetology 64:18–38.

2015. Furman*, B. L. S., A. J. Bewick*, T. L. Harrison**, E. Greenbaum, V. Gvoždík, C. Kusamba, and B. Evans. Pan-African phylogeography of a model organism, the African clawed frog “Xenopus laevis.” Molecular Ecology 24:909–925.

2015. Portillo*, F., E. Greenbaum, M. Menegon, C. Kusamba, and J. M. Dehling. Phylogeography and species boundaries of Leptopelis (Anura: Arthroleptidae) from the Albertine Rift. Molecular Phylogenetics and Evolution 82:75–86.

2014. Greenbaum, E., J. Meece, K. Reed, and C. Kusamba. Amphibian chytrid infections in non-forested habitats of Katanga, Democratic Republic of the Congo. Herpetological Review 45:610–614.

2014. Leaché, A. D., P. Wagner, C. W. Linkem, W. Böhme, T. J. Papenfuss, R. Chong, B. R. Lavin**, A. M. Bauer, S. Nielsen*, E. Greenbaum, M. -O. Rödel, A. Schmitz, M. LeBreton, I. Ineich, L. Chirio, E. A. Eniang, S. Baha El Din, A. R. Lemmon, and F. T. Burbrink. A hybrid phylogenetic-phylogenomic approach for species tree estimation in African Agama lizards with applications to biogeography, character evolution, and diversification. Molecular Phylogenetics and Evolution 79:215–230.

2014. Portillo*, F., and E. Greenbaum. A new species of the Leptopelis modestus complex (Anura: Arthroleptidae) from the Albertine Rift of Central Africa. Journal of Herpetology 48:394–406.

2014. Wagner, P., E. Greenbaum, P. Malonza, and W. R. Branch. Resolving sky island speciation in populations of East African Adolfus alleni (Sauria, Lacertidae). Salamandra 50:1–17.

2014. Heinicke, M. P., Juan D. Daza, E. Greenbaum, T. R. Jackman, and A. M. Bauer. Phylogeny, taxonomy, and biogeography of a circum-Indian Ocean clade of leaf-toed geckos (Reptilia: Gekkota), with a description of two new genera. Systematics and Biodiversity 12:23–42. Awarded outstanding paper in 2014 by journal's editorial board.

2014. Portillo*, F., and E. Greenbaum. At the edge of a species boundary: A new and relatively young species of Leptopelis (Anura: Arthroleptidae) from the Itombwe Plateau, Democratic Republic of the Congo. Herpetologica 70:100–119.

2013. Greenbaum, E., U. Sinsch, E. Lehr, F. Valdez**, and C. Kusamba. Phylogeography of the reed frog Hyperolius castaneus (Anura: Hyperoliidae) from the Albertine Rift of Central Africa: Implications for taxonomy, biogeography and conservation. Zootaxa 3731:473–494.

2013. Böhm, M., B. Collen, J. E. M. Baillie, J. Chanson, N. Cox, G. Hammerson, M. Hoffmann, S. R. Livingstone, M. Ram, A. G. J. Rhodin, S. N. Stuart, P. P. van Dijk, B. Young, L. E. Afuang, A. Aghasyan, A. G. Aguayo, C. Aguilar, R. Ajtic, F. Akarsu, L. R. V. Alencar, A. Allison, N. Ananjeva, S. Anderson, C. Andrén, D. Ariano-Sánchez, J. C. Arredondo, M. Auliya, C. C. Austin, A. Avci, P. J. Baker, A. F. Barreto-Lima, C. L. Barrio-Amorós, D. Basu, M. F. Bates, A. Batistella, A. Bauer, D. Bennett, W. Böhme, D. Broadley, R. Brown, J. Burgess, A. Captain, S. Carreira, M. R. Castaneda, F. Castro, A. Catenazzi, J. R. Cedeño-Vázquez, D. Chapple, M. Cheylan, D. F. Cisneros-Heredia, D. Cogalniceanu, H. Cogger, C. Corti, G. C. Costa, P. J. Couper, T. Courtney, J. Crnobrnja-Isailovic, P. -A.Crochet, B. Crother, F. Cruz, J. Daltry, R. J. R. Daniels, I. Das, A. de Silva, A. C. Diesmos, L. Dirksen, T. M. Doan, K. Dodd, J. S. Doody, M. E. Dorcas, J. Duarte de Barros Filho, V. T. Egan, E. H. El Mouden, D. Embert, R. E. Espinoza, A. Fallabrino, X. Feng, Z. -J. Feng, L. Fitzgerald, O. Flores-Villela, F. G. R. França, D. Frost, H. Gadsden, T. Gamble, S. R. Ganesh, M. A. Garcia, J. E. García-Pérez, J. Gatus, M. Gaulke , P. Geniez, A. Georges, J. Gerlach, S. Goldberg, J. -C. T. Gonzalez, D. J. Gower, T. Grant, E. Greenbaum, C. Grieco, P. Guo, A. M. Hamilton, G. Hammerson, K. Hare, B. Hedges, N. Heideman, C. Hilton-Taylor, R. Hitchmough, M. Hoffmann, B. Hollingsworth, M. Hutchinson, I. Ineich, J. Iverson, F. M. Jaksic, R. Jenkins, U. Joger, R. Jose, Y. Kaska, U. Kaya, J. S. Keogh, G. Köhler, G. Kuchling, Y. Kumlutaş, A. Kwet, E. La Marca, W. Lamar, A. Lane, B. Lardner, C. Latta, G. Latta, M. Lau, P. Lavin, D. Lawson, M. LeBreton, E. Lehr, D. Limpus, N. Lipczynski, S. Livingstone, A. S. Lobo, M. A. López-Luna, L. Luiselli, V. Lukoschek, M. Lundberg, P. Lymberakis, R. Macey, W. E. Magnusson, D. L. Mahler, A. Malhotra, J. Mariaux, B. Maritz, O. A. V. Marques, R. Márquez, M. Martins, G. Masterson, J. A. Mateo, R. Mathew, N. Mathews, G. Mayer, J. R. McCranie, G. J. Measey, F. Mendoza-Quijano, M. Menegon, S. Métrailler, D. A. Milton, C. Montgomery, S. A. A. Morato, T. Mott, A. Muñoz-Alonso, J. Murphy, T. Q. Nguyen, G. Nilson, C. Nogueira, H. Nunez, N. Orlov, H. Ota, J. Ottenwalder, T. Papenfuss, S. Pasachnik, P. Passos, O. S. G. Pauwels, N. Pérez-Buitrago, V. Pérez-Mellado, E. R. Pianka, J. Pleguezuelos, C. Pollock, P. Ponce-Campos, R. Powell, F. Pupin, G. E. Quintero Díaz, R. Radder, J. Ramer, A. R. Rasmussen, C. Raxworthy, R. Reynolds, N. Richman, E. L. Rico, E. Riservato, G. Rivas, P. L. B. Rocha, M. -O. Rödel, L. Rodríguez Schettino, W. M. Roosenburg, R. Sadek, K. Sanders, G. Santos-Barrera, H. H. Schleich, B. R. Schmidt, A. Schmitz, M. Sharifi, G. Shea, H. Shi, R. Shine, R. Sindaco, T. Slimani, R. Somaweera, P. Stafford, R. Stuebing, S. Sweet, E. Sy, H. Temple, M. F. Tognielli, K. Tolley, P. J. Tolson, B. Tuniyev, S. Tuniyev, N. Üzüm, G. van Buurt, M. Van Sluys, A. Velasco, M. Vences, M. Veselý, S. Vinke, T. Vinke, G. Vogel, M. Vogrin, R. C. Vogt, O. R. Wearn, Y. L. Werner, M. J. Whiting, T. Wiewandt, J. Wilkinson, B. Wilson, S. Wren, T. Zamin, K. Zhou, and G. Zug. Conservation status of the world’s reptiles. Biological Conservation 157:372–385.

2012. Heinicke, M. P., E. Greenbaum, T. R. Jackman, and A. M. Bauer. Evolution of gliding in Southeast Asian geckos and other vertebrates is temporally congruent with dipterocarp forest development. Biology Letters 8:994–997.

2012. de Sá, R. O., J. W. Streicher, R. Selenkoyela, M. C. Forlani, S. P. Loader, E. Greenbaum, S. Richards, and C. F. B. Haddad. Molecular phylogeny of microhylid frogs (Anura: Microhylidae) with emphasis on relationships among New World genera. BMC Evolutionary Biology 12:241 (21 pp).

2012. Anderson*, C. G., and E. Greenbaum. Phylogeography of northern populations of the black-tailed rattlesnake (Crotalus molossus Baird and Girard, 1853), with the revalidation of C. ornatus Hallowell, 1854. Herpetological Monographs 26:19–57.

2012. Greenbaum, E., F. Portillo*, and C. Kusamba. Systematics of the poorly known treefrog Leptopelis fiziensis (Anura: Arthroleptidae), with a description of its call. African Journal of Herpetology 61:113–127.

2012. Greenbaum, E., and C. Kusamba. Conservation implications following the rediscovery of four frog species from the Itombwe Natural Reserve, eastern Democratic Republic of the Congo. Herpetological Review 43:253–259.

2012. Gamble, T., E. Greenbaum, T. R. Jackman, A. P. Russell, and A. M. Bauer. Repeated origin and loss of adhesive toepads in geckos. PLoS ONE 7:e39429. Article of special significance by Faculty of 1000;

2012. Wagner, P., E. Greenbaum, and A. M. Bauer. A new species of the Acanthocercus atricollis (Squamata: Agamidae) complex from Zambia. Salamandra 48:21–30.

2012. Greenbaum, E., E. L. Stanley*, C. Kusamba, W. M. Moninga, S. R. Goldberg, and C. R. Bursey. A new species of Cordylus (Squamata: Cordylidae) from the Marungu Plateau of south-eastern Democratic Republic of the Congo. African Journal of Herpetology 61:14–39.

2012. Oliver, P. M., A. M. Bauer, E. Greenbaum, T. Jackman, and T. Hobbie**. Molecular phylogenetics of the arboreal Australian gecko genus Oedura Gray 1842 (Gekkota: Diplodactylidae): Another plesiomorphic grade? Molecular Phylogenetics and Evolution 63:255–264.

2012. Greenbaum, E., K. A. Tolley, A. Joma*, and C. Kusamba. A new species of chameleon (Sauria: Chamaeleonidae: Kinyongia) from the northern Albertine Rift, Central Africa. Herpetologica 68:60–75.

2011. Sinsch, U., E. Greenbaum, C. Kusamba and E. Lehr. Rapid assessment of montane anuran communities in the Albertine Rift: Hyperolius castaneus Ahl, 1931 as a focal species for conservation. African Zoology 46:320–333.

2011. Greenbaum, E., C. O. Villanueva**, C. Kusamba, M. M. Aristote and W. R. Branch. A molecular phylogeny of Equatorial African Lacertidae, with the description of a new genus and species from eastern Democratic Republic of the Congo. Zoological Journal of the Linnean Society 163:913–942.

2011. Heinicke, M. P., E. Greenbaum, T. Jackman, and A. M. Bauer. Phylogeny of a trans-Wallacean radiation (Squamata, Gekkonidae, Gehyra) supports a single early colonization of Australia. Zoologica Scripta 40:584–602.

2011. Roelke*, C. E., E. Greenbaum, C. Kusamba, M. M. Aristote, and E. N. Smith. Systematics and conservation status of two distinct Albertine Rift treefrogs, Leptopelis karissimbensis and Leptopelis kivuensis (Anura: Arthroleptidae). Journal of Herpetology 45:343–351.

2011. Greenbaum, E., E. N. Smith, and R. O. de Sá. Molecular systematics of the Middle American genus Hypopachus (Anura: Microhylidae). Molecular Phylogenetics and Evolution 61:265–277.

2011. Rösler, H., A. M. Bauer, M. Heinicke, E. Greenbaum, T. Jackman, T. Q. Nguyen, and T. Ziegler. Phylogeny, taxonomy and zoogeography of the genus Gekko Laurenti, 1768 with the revalidation of G. reevesii Gray, 1831 (Sauria: Gekkonidae). Zootaxa 2989:1–50.

2011. Gamble, T., A. M. Bauer, G. R. Colli, E. Greenbaum, L. J. Vitt, and A. M. Simons. Coming to America: Multiple origins of New World geckos. Journal of Evolutionary Biology 24:231–244.

2011. Evans, B. J., E. Greenbaum, C. Kusamba, T. F. Carter, M. L. Tobias**, S. A. Mendel**, and D. B. Kelley. Description of a new octoploid frog species (Anura: Pipidae: Xenopus) from the Democratic Republic of the Congo, with a discussion of the biogeography of African clawed frogs in the Albertine Rift. Journal of Zoology, London 283:276–290.

2010. Heinicke, M. P., E. Greenbaum, T. R. Jackman, and A. M. Bauer. Molecular phylogenetics of Pacific Nactus (Squamata: Gekkota: Gekkonidae) and the diphyly of Australian species. Proceedings of the California Academy of Sciences 61:633–646.

2010. Bauer, A. M., T. R. Jackman, E. Greenbaum, A. de Silva, V. B. Giri, and I. Das. Molecular evidence for the taxonomic status of Hemidactylus brookii group taxa (Squamata: Gekkonidae). Herpetological Journal 20:129–138.

2010. Bauer, A. M., T. R. Jackman, E. Greenbaum, V. B. Giri, and A. de Silva. South Asia supports a major endemic radiation of Hemidactylus geckos. Molecular Phylogenetics and Evolution 57:343–352.

2008. Bauer, A. M., V. B. Giri, E. Greenbaum, T. R. Jackman, M. S. Dharne, and Y. S. Shouche. On the systematics of the gekkonid genus Teratolepis Günther, 1869: another one bites the dust. Hamadryad 33:13–27.

2008. Jackman, T. R., A. M. Bauer, and E. Greenbaum. Phylogenetic relationships of geckos of the genus Nactus and their relatives (Squamata: Gekkonidae). Acta Herpetologica 3:1–18.

2008. Simmons, J. E., A. C. Campbell, and E. Greenbaum. Presence of elemental mercury in fluid preserved specimens. Collection Forum 22:32–36.

2008. McCranie, J. R., and E. Greenbaum. Bolitoglossa heiroreias Greenbaum. Catalogue of American Amphibians and Reptiles 842:1–3.

2008. Gamble, T., A. M. Bauer, E. Greenbaum, and T. R. Jackman. Out of the blue: a novel, trans-Atlantic clade of gecko lizards (Gekkota, Squamata). Zoologica Scripta 37:355–366.

2008. Greenbaum, E., C. Kusamba, M. Aristote, and K. D. Reed. Amphibian chytrid fungus infections in Hyperolius (Anura: Hyperoliidae) from eastern Democratic Republic of Congo. Herpetological Review 39:70–73.

2008. Gamble, T., A. M. Bauer, E. Greenbaum, and T. R. Jackman. Evidence for Gondwanan vicariance in an ancient clade of gecko lizards. Journal of Biogeography 35:88–104.

2007. Greenbaum, E., A. M. Bauer, and T. R. Jackman. Homopholis and Blaesodactylus (Squamata: Gekkonidae) revisited: new insights from a molecular phylogeny. African Journal of Herpetology 56:101–114.

2007. Jackman, T. R., A. M. Bauer, E. Greenbaum, M. Vences, and F. Glaw. Molecular phylogenetic relationships among species of the Malagasy-Comoran gecko genus Paroedura (Sauria: Gekkonidae). Molecular Phylogenetics and Evolution 46:74–81.

2007. Greenbaum, E., A. M. Bauer, T. R. Jackman, M. Vences, and F. Glaw. A phylogeny of the enigmatic Madagascan geckos of the genus Uroplatus (Sauria: Gekkonidae). Zootaxa 1493:41–51.

2007. Pramuk, J. B., M. J. Grose, A. L. Clarke, E. Greenbaum, E. Bonaccorso, J. M. Guayasamin, A. H. Smith, B. W. Benz, B. R. Harris, E. Siegfreid, Y. R. Reid, N. Holcroft, and E. O. Wiley. Phylogeny of finescale shiners of the genus Lythrurus (Cypriniformes: Cyprinidae) inferred from four mitochondrial genes. Molecular Phylogenetics and Evolution 42:287–297.

2007. Bauer, A. M., T. R. Jackman, E. Greenbaum, and T. J. Papenfuss. First record of Lepidodactylus lugubris in Suriname. Applied Herpetology 4:84–85.

2007. Herrera, N., V. Henríquez, and E. Greenbaum*. New department and country records for amphibians and reptiles from El Salvador. Herpetological Review 38:222–226. *Corresponding author

2007. Bauer, A. M., A. De Silva, E. Greenbaum, and T. R. Jackman. A new species of day gecko from high elevation in Sri Lanka, with a preliminary phylogeny of Sri Lankan Cnemaspis (Reptilia: Squamata: Gekkonidae). Mitteilungen aus dem Museum für Naturkunde in Berlin. Zoologische Reihe 83 (Sonderheft):22–32.

2006. Bauer, A. M., T. Jackman, E. Greenbaum, and T. J. Papenfuss. Confirmation of the occurrence of Hemidactylus robustus Heyden, 1827 (Reptilia: Gekkonidae) in Iran and Pakistan. Zoology in the Middle East 39:59–62.

2006. Greenbaum, E., A. C. Campbell, and C. J. Raxworthy. A revision of sub-Saharan Chalcides (Squamata: Scincidae), with redescriptions of two East African species. Herpetologica 62:71–89.

2005. Greenbaum, E., and O. Komar. Threat assessment and conservation prioritization of the herpetofauna of El Salvador. Biodiversity and Conservation 14:2377–2395.

2005. Greenbaum, E. Systematics of West African skinks in the Chalcides thierryi group: composition, distribution, and redescription of types. African Journal of Herpetology 54:17–29.

2005. Greenbaum, E., and J. L. Carr. The herpetofauna of Upper Niger National Park, Guinea, West Africa. Scientific Papers, Natural History Museum, The University of Kansas 37:1–21.

2005. Sheil, C. A., and E. Greenbaum. Reconsideration of skeletal development of Chelydra serpentina (Reptilia: Testudinata: Chelydridae): evidence for intraspecific variation. Journal of Zoology, London 265:235–267.

2004. Greenbaum, E., J. L. Carr, and A. Almendáriz. Taxonomic status of Tantilla equatoriana Wilson and Mena 1980 (Serpentes: Colubridae). The Southwestern Naturalist 49:457–464.

2004. Greenbaum, E. A new species of Bolitoglossa (Amphibia: Caudata: Plethodontidae) from montane forests in Guatemala and El Salvador. Journal of Herpetology 38:411–421.

2004. Greenbaum, E., and M. Jorgensen. Envenomated-invertebrate prey preference of Agkistrodon contortrix (Serpentes: Viperidae) during strike-induced chemosensory searching. Amphibia-Reptilia 25:165–172.

2004. Greenbaum, E. The influence of prey-scent stimuli on predatory behavior of the North American copperhead Agkistrodon contortrix (Serpentes: Viperidae). Behavioral Ecology 15:345–350.

2003. Greenbaum, E., N. Galeva, and M. Jorgensen. Venom variation and chemoreception of the viperid Agkistrodon contortrix: evidence for adaptation? Journal of Chemical Ecology 29:1741–1755.

2002. Greenbaum, E. A standardized series of embryonic stages for the emydid turtle Trachemys scripta. Canadian Journal of Zoology 80:1150–1170.

2002. Greenbaum, E., and J. L. Carr. Staging criteria for embryos of the spiny softshell turtle, Apalone spinifera (Testudines: Trionychidae). Journal of Morphology 254:272–291.

2001. Greenbaum, E., and J. L. Carr. Sexual differentiation in the spiny softshell turtle (Apalone spinifera), a species with genetic sex determination. Journal of Experimental Zoology 290:190–200.

2010. Greenbaum, E., and O. Komar. A conservation assessment of Salvadoran protected areas: priorities and recommendations based on amphibian and reptile distributions, pp. 436–459 In: Wilson, L. D., J. H. Townsend, and J. D. Johnson (eds.), Conservation of Mesoamerican Amphibians and Reptiles. Eagle Mountain Publishing, LC, Eagle Mountain, Utah.

2008. Bauer, A. M., T. R. Jackman, E. Greenbaum, and T. Gamble. Phylogenetic relationships of the Italian gekkotan fauna, pp. 59–62 In: Corti, C. (ed.), Herpetologia Sardiniae. Societas Herpetologica Italica, Edizioni Belvedere, Latina, “le scienze” (8), 504 pp.

2017.   Greenbaum, E. Emerald Labyrinth: A Scientist’s Adventures in the Jungles of the CongoForeEdge (imprint of University Press of New England). xv + 329 pp.

2006. Köhler, G., M. Veselý, and E. Greenbaum*The Amphibians and Reptiles of El Salvador. Krieger Publishing Company, Malabar, FL. ix + 238 pp. *Corresponding author

2020. Greenbaum, E. 野性与文明一位进化遗传学家在刚果(金)丛林的新发现 [Chinese translation of Emerald Labyrinth by Zhu Hongfei]. Grand China Publishing House, Beijing, China. 301 pp.

____.  Coto, M., V. Henríquez, and E. Greenbaum. Geographic Distribution. Bothriechis thalassinus (Merendon Palm-Pit Viper). Herpetological Review In press. 8 May 2023.

2022.  Goldberg, S., C. Bursey, and E. Greenbaum. Endoparasites. Hemidactylus mabouia (House Gecko). Herpetological Review 53:680. 

2022. Blackburn, D. C., V. Dérozier, A. Djaman, E. Greenbaum, and G. F. M. Jongsma. First record of Spiny Frogs, Acanthixalus (Hyperoliidae, Kassininae), from south of the Congo River. Herpetology Notes 15:641–647.

2021. Goldberg, S. R., C. R. Bursey, C. Kusamba, and E. Greenbaum. Endoparasites. Leptopelis calcaratus (Spurred Forest Treefrog). Herpetological Review 52:830–831.

2021. Goldberg, S. R., C. R. Bursey, C. Kusamba, and E. Greenbaum. Endoparasites. Atheris nitschei (Great Lakes Bush Viper). Herpetological Review 52:657.

2021. Goldberg, S. R., C. R. Bursey, C. Kusamba, and E. Greenbaum. Endoparasites. Philothamnus angolensis (Angola Green Snake). Herpetological Review 52:675.

2021. Greenbaum, E., D. F. Hughes, and M. Behangana. Geographic Distribution. Leptopelis calcaratus (Spurred Forest Treefrog). Herpetological Review 52:342.

2018. Hughes*, D. F., M. Behangana, L. Wilber, and E. Greenbaum. Amietia nutti (Nutt’s River Frog): Leech parasitism. African Herp News (69)15–19.

2017.   Greenbaum, E., C. Kusamba, W. M. Muninga, and M. M. Aristote. Geographic Distribution. Boaedon perisilvestris (Near-Forest House Snake). Herpetological Review 48:127.

2016. Greenbaum, E., C. Kusamba, and M. Luhumyo. Geographic Distribution. Hyperolius rwandae (Rwanda Long Reed Frog). Herpetological Review 47:419.

2016. Hughes*, D. F., E. Greenbaum, and M. Behangana. Diet and Feeding Behavior. Lycophidion ornatum (Ornate Wolf Snake). Herpetological Review 47:147–148.

2015. Morán**, E. S., V. Henríquez, E. Greenbaum, J. G. C. López, and A. M. Rivera. Geographic Distribution. Sibon dimidiatus (Slender Snail Sucker). Herpetological Review 46:576.

2015. Greenbaum, E., D. F. Hughes*, C. Kusamba, and F. M. Masudi**. Geographic Distribution. Leptopelis mackayi (Mackay’s Forest Treefrog). Herpetological Review 46:378.

2014. Goldberg, S. R., C. R. Bursey, E. Greenbaum, C. Kusamba, and J. Arreola**. Gerrhosauridae. Gerrhosaurus nigrolineatus Hallowell, 1857. Black-lined Plated Lizard. Endoparasites. African Herp News 61:20–21.

2014. Henríquez, V., and E. Greenbaum. Geographic Distribution. Hyalinobatrachium fleischmanni (Fleischmann’s Glass Frog). Herpetological Review 45:87.

2013. Henríquez, V., and E. Greenbaum. Geographic Distribution. Bolitoglossa salvinii (Salvin's Mushroomtongue Salamander). Herpetological Review 44:470.

2012. Goldberg, S. R., C. R. Bursey, E. Greenbaum, and C. Kusamba. Gastrointestinal helminths of the black-necked agama, Acanthocercus atricollis (Squamata: Agamidae), from Democratic Republic of the Congo. Comparative Parasitology 79:164–166.

2012. Greenbaum, E., C. Kusamba, W. Moninga, and M. M. Aristote. Hemisus marmoratus (Marbled Snout-Burrower). Defensive Behavior. Herpetological Review 43:320–321.

2010. Greenbaum, E., K. Jackson, and C. Kusamba. Colubridae: Natriciteres olivacea (Peters, 1854): Olive Marsh Snake. Melanism. African Herp News (52):10–11.

2010. Greenbaum, E., and C. Kusamba. Geographic Distribution. Xenopus ruwenzoriensis (Ruwenzori Clawed Frog). Herpetological Review 41:376–377.

2009. Ibarra Portillo, R., V. Henríquez, and E. Greenbaum. Geographic Distribution. Trachemys emolli (Moll’s Slider). Herpetological Review 40:111.

2008. Greenbaum, E. Book review. Biology of Turtles. By Jeanette Wyneken, Matthew H. Godfrey, and Vincent Bels. Herpetological Review 39:376–378.

2007. Gamble, T., and E. Greenbaum. Geographic Distribution. Aspidoscelis sexlineatus (Six-lined Racerunner). Herpetological Review 38:482.

2007. Greenbaum, E. Book review. Reptiles and Amphibians of East Africa. By Stephen Spawls, Kim M. Howell, and Robert C. Drewes. Herpetological Review 38:121–123.

2006. Greenbaum, E., and D. M. Davis. Geographic Distribution. Storeria dekayi wrightorum (Midland brown snake). Herpetological Review 37:109.

2006. Greenbaum, E. Geographic Distribution. Tropidoclonion lineatum (Lined snake). Herpetological Review 37:110.

2005. Greenbaum, E. Book review. Amphibians and Reptiles of the Bay Islands and Cayos Cochinos, Honduras. By James R. McCranie, Larry David Wilson, and Gunther Köhler. Herpetological Review 36:483–484.

2004. Greenbaum, E., and J. B. Rasmussen. Chalcides thierryi pulchellus (Cylindrical skink). Attempted predation. Herpetological Review 35:166–167.

2003. Greenbaum, E., M. C. Baldé, and J. L. Carr. Geographic Distribution. Dendroaspis polylepis (Black mamba). Herpetological Review 34:168.

2002. Greenbaum, E., N. Herrera, R. Ibarra Portillo, O. Komar, and R. Rivera. Geographic Distribution. Mesoscincus managuae (Managua skink). Herpetological Review 33:322.

2002. Greenbaum, E., O. Komar, and R. Bolaños. Geographic Distribution. Ninia espinali (Espinal's coffee snake). Herpetological Review 33:325.

2002. Greenbaum, E. Geographic Distribution. Hemidactylus frenatus (Common house gecko). Herpetological Review 33:65–66.

2000. Greenbaum, E. Geographic Distribution. Hyla avivoca (Bird-voiced treefrog). Herpetological Review 31:251.

2009. Greenbaum, E., and M. J. Whiting. Homopholis walbergii. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.4. <>.

2005. Greenbaum, E., and T. Leenders. Bolitoglossa heiroreias. AmphibiaWeb: Information on amphibian biology and conservation. [web application]. 2005. Berkeley, California: AmphibiaWeb. Available:

2017. Hughes, D. F., E. Greenbaum, L. Wilber, and M. Behangana. Expedition Report from Uganda: Herpetological Surveys at Eight Central Forest Reserves (Agoro-Agu, Budongo, Echuya, Mabira, Mount Kadam, Mount Kei, Mount Moroto, and Mount Otze) and Four National Parks (Bwindi Impenetrable, Mgahinga Gorilla, Mount Elgon, and Rwenzori Mountains). Report to Uganda Wildlife Authority available at:

2015. Seimon, T., S. Ayebare, D. Meirte, D. Kujirakwinja, G. Mitamba, E. Muhindo, D. McAloose, M. Behangana, W. Lukwago, E. Greenbaum, and A. Plumptre. FINAL REPORT: US Fish and Wildlife Service (Grant no. F13AP00871). Modeling Disease Risk in Threatened Amphibians in the Albertine Rift.

2014. Hughes, D. F., E. Greenbaum, and M. Behangana.  Expedition Report from Uganda: Herpetological Surveys at Bwindi Impenetrable National Park, Rwenzori Mountains National Park, and the northern shore of Lake Victoria.  Report to Uganda Wildlife Authority available at:

2010. Greenbaum, E. FINAL REPORT: National Geographic Research and Exploration Grant (no. 8556-08), $18,522. Conservation Assessment of Endangered Amphibians of the Itombwe Highlands, Democratic Republic of Congo. 11 pp.

Invited Web Publications (2 Publications)

Greenbaum, E. 2017. When Science Means Getting Cobra Venom Spat Into Your Eye.

Greenbaum, E. 2017. The Itombwe Golden Frog’s Uncertain Future. Society for Conservation Biology, Africa Section.

2018.  Greenbaum, E. Congo quest for cobras. Reptiles Magazine 26:32–39.

2008. Greenbaum, E. The fragile forests. Africa Geographic 16:51–57.

2003. Greenbaum, E. A practical and safe method for the capture and restraint of venomous snakes in the field. Bulletin of the Chicago Herpetological Society 38:1–3.

2000. Greenbaum, E. Herpetofaunal observations in disparate habitats in South Australia, New South Wales, and Queensland, Australia. Herpetological Bulletin 72:6–16.

1999. Greenbaum, E. The enigma of sex determination in turtles. Bulletin of the Chicago Herpetological Society 34:113–115.

1999. Greenbaum, E. Efficiency of three amphibian and reptile population estimation methods used at D'Arbonne National Wildlife Refuge, Union parish, Louisiana. Bulletin of the Chicago Herpetological Society 33:191–193.

1988. Greenbaum, E. Snakes under pressure. Notes from NOAH 16:9–10.

1986. Greenbaum, E. Serpentarium adventure. Notes from NOAH 13:14–15.

2004. Greenbaum, E., C. Sheil, D. McLeod, and B. Beatty. Supplemental Materials: Human Anatomy Observation (BIOL 301) & Dissection (BIOL 302) Laboratories. The University of Kansas, Lawrence, KS. 62 pp.